Mysterious objects thought to be meteorites crash into Earth's oceans near many major cities. These objects are revealed to be spacecraft containing hostile extraterrestrial beings. U.S. Marines from Camp Pendleton arrive in LA to defend against alien ground forces and assist in the evacuation of civilians in preparation for a bombing campaign. Among the military's forces is SSgt. Michael Nantz (Aaron Eckhart), a 20-year veteran who lost his squad during his last deployment in Iraq. Nantz had planned to retire, but the situation requires him to replace the platoon sergeant from Echo Company, 2nd Battalion 5th Marines.
Under command of 2ndLt. William Martinez (Ramon Rodriguez), fresh out of The Basic School, the platoon attends a briefing at Santa Monica Airport, which has been converted to a Forward Operating Base. They learn that the alien force consists solely of ground forces at the moment, with no air support to speak of; as such, the Air Force plans to commence saturation bombing of the Santa Monica area at 1930 hours. The platoon's mission is to rescue civilians from an LAPD police station within three hours. As they advance through the city, they are ambushed and suffer casualties, but link up with a group of Army National Guard soldiers from the 40th Infantry Division, including Air Force intelligence Technical Sergeant Elena Santos (Michelle Rodriguez). Eventually, the combined team reaches the police station, finding five civilians inside: a veterinarian named Michele (Bridget Moynahan), children Hector Rincon (Bryce Cass), Kirsten (Joey King) and Amy (Jadin Gould), and Hector's father Joe (Michael Peña). A helicopter arrives to evacuate the wounded Marines Simmons, Guerrero, Lenihan and Grayston, but is unable to take on the civilians because it is already full. As the helicopter is taking off, it is destroyed by an alien aircraft, killing everyone aboard.
The Marines commandeer a city bus for evacuation. They also find a wounded alien and, with the help of Michele, learn of the aliens' physiology, finding a weak point in the torso. They also learn that the alien aircraft are remote-controlled drones and that the drones can track human radio transmissions. Santos reveals her original mission was to locate the aliens' central command center, believing that its destruction would deactivate the alien drones and turn the tide of the war. On a freeway, they encounter a lone infantry fighting vehicle that is under attack and subsequently destroyed by the aliens. With the bus under attack and the freeway off-ramp destroyed, the Marines begin rappelling the civilians off the side of the freeway. The Army personnel, and Stravou and Mottola are killed. Mr. Rincon and Lt. Martinez are also wounded. Martinez sacrifices himself to destroy the enemy unit by detonating C-4 explosives inside the bus, leaving Nantz in command.
The remnants of the platoon (Nantz, Santos, Corporal Kevin Harris (Ne-Yo), Corpsman Jibril Adukwu (Adetokumboh M'Cormack), Lance Corporal Peter Kerns (Jim Parrack), Corporal Lee Imlay (Will Rothhaar), Corporal Jason Lockett (Cory Hardrict) and the five civilians) escape the bombing zone, taking refuge at a convenience store. A news report speculates that the aliens have attacked Earth for its water supply, which they use as fuel. The team waits for the bombing to begin, but nothing happens. The reason for this is revealed at the Santa Monica Airport, where the FOB was destroyed by the aliens, and Nantz learns that the military forces are losing Los Angeles. Utilizing an LAV-25 and a Humvee, Echo Company plans to escort the civilians to a helicopter at an alternate extraction point. Before they embark, Rincon dies from his wounds and Lockett (whose brother died under Nantz's command in Iraq) openly doubts Nantz's ability to lead. Nantz and Lockett come to terms with their guilt and anger as Nantz lists the names, ranks and serial numbers of all the marines who have died under his command from memory. They reach the extraction point and leave.
From the air, Nantz spots a city block lacking power, and speculates that the block is the location of the alien command center. With the military forces having retreated from the area, Nantz decides to try to recon the area alone, but his team insists on accompanying him. Going underground, the Marines confirm the presence of a large alien vessel and return to the surface. Kerns radios in to request guided missile strikes, which will be targeted by Nantz via laser designator while the remaining Marines defend the position. Kerns is killed when his radio broadcasts draw attention to him, and the command center, damaged but not disabled by the first missile, rises from the ground to find safer haven. The drones protect their command center from the second missile, but the marines use an AT4 rocket launcher to destroy the drone that was about to intercept the third missile. The third missile strikes the command center, which soon falls apart. The alien ground forces retreat as their drones rain down around them, and reinforcements arrive to extract the surviving marines in Los Angeles.
Nantz's team is extracted to a temporary base in the Mojave Desert, where they are greeted as heroes for their bravery. All other cities under attack now plan to emulate their strategy, destroying the alien command center in each city. Despite orders to rest, Nantz's team rejoin the rest of the Marines in retaking Los Angeles
Battle:Los Angeles
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